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Similar content being viewed by others Introduction Nicotiana benthamiana has become a model plant for studying plant-pathogen interactions 1,2 due to its unusual susceptibility to a wide range of viruses 3 and has developed important critical tools such as virus-induced gene silencing (VIGS) 4,5 One of the essential reasons for plant susceptibility is the R gene, which is responsible for pathogen detection and host defense in plants and the majority of them encode the nucleotide-binding site leucine-rich repeat (NBS-LRR) proteins 6

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